1
: the wedge-shaped piece at the crown of an arch that locks the other pieces in place see arch illustration
2
: something on which associated things depend for support
determination, a keystone of the puritan ethic—L. S. Lewis
3
or keystone species
: a species of plant or animal that produces a major impact (as by predation) on its ecosystem and is considered essential to maintaining optimum ecosystem function or structure
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Merriam-Webster unabridged
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