anucleolate
adjective
anu·cle·o·late
(ˈ)ā-n(y)ü-ˈklē-ə-ˌlāt
-lət,
-ˈn(y)ü-klē-ə-
: lacking a cell nucleolus
These findings were corroborated by the anucleolate mutant, which served as a control, since it lacks ribosomal RNA genes.—Walter J. Gehring, Master Control Genes in Development and Evolution … , 1998
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Merriam-Webster unabridged
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